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Table 1 Listing of Heteroscleromorpha genera in alphabetical order

From: Proposal for a revised classification of the Demospongiae (Porifera)

Genus

Order

Family

New order (Suborder)

Family

Genes, type species

Genes, non-type

Aaptos

Hadromerida

Suberitidae

Suberitida

Suberitidae

no

yes

Abyssocladia

Poecilosclerida

Cladorhizidae

Poecilosclerida

Cladorhizidae

no

yes

Acalle

Haplosclerida

Metaniidae

Spongillida

Metaniidae

no

no

Acanthancora

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

18S

no

Acanthella

Halichondrida

Dictyonellidae

Bubarida

Dictyonellidae

18S, 28S, COI

yes

Acantheurypon

Poecilosclerida

Raspailiidae

Poecilosclerida

Poecilosclerida incertae sedis

18S, 28S, COI

yes

Acanthochaetetes

Hadromerida

Acanthochaetetidae

Clionaida

Acanthochaetetidae

no

yes

Acanthoclada

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Acanthopolymastia

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Acanthorhabdus

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Acanthostylotella

Poecilosclerida

Raspailiidae

Agelasida

Agelasidae

18S, 28S

no

Acanthotetilla

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

no

yes

Acanthotriaena

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

Pachastrellidae

no

no

Acanthotylotra

Haplosclerida

Spongillina incertae sedis

Spongillida

Spongillida incertae sedis

no

no

Acarnus

Poecilosclerida

Acarnidae

Poecilosclerida

AcarnidaeCP

16S, 18S, 28S

no

Acheliderma

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Aciculites

‘Lithistida’

Scleritodermidae

Tetractinellida (Spirophorina)

Scleritodermidae

no

yes

Adreus

Hadromerida

Hemiasterellidae

Tethyida

Hemiasterellidae

18S, 28S, COI

yes

Agelas

Agelasida

Agelasidae

Agelasida

Agelasidae

28S, COI

yes

Alectona

Astrophorida

Thoosidae

Tetractinellida (Astrophorina)

Thoosidae

COI, 28S

no

Alloscleria

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Amorphinopsis

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

18S, 28S, COI, EF-1

yes

Amphiastrella

Poecilosclerida

Iotrochotidae

Poecilosclerida

Iotrochotidae

no

no

Amphibleptula

‘Lithistida’

Scleritodermidae

Tetractinellida (Spirophorina)

Scleritodermidae

no

no

Amphilectus

Poecilosclerida

Esperiopsidae

Poecilosclerida

Esperiopsidae

no

yes

Amphinomia

Poecilosclerida

Echinodictyinae

Poecilosclerida

Echinodictyinae

18S, 28S

no

Amphitethya

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

no

yes

Anaderma

‘Lithistida’

Pleromidae

Tetractinellida (Astrophorina)

Pleromidae

no

no

Ancorella

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

Pachastrellidae

no

no

Ancorina

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

yes

Anheteromeyenia

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Anisocrella

Poecilosclerida

Crellidae

Poecilosclerida

Crellidae

no

no

Annulastrella

Astrophorida

Theneidae

Tetractinellida (Astrophorina)

Theneidae

no

yes

Antho

Poecilosclerida

Ophlitaspongiinae

Poecilosclerida

Microcionidae

18S, 28S

yes

Anthotethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Arabescula

‘Lithistida’

Lithistida incertae sedis

incertae sedis

 

no

no

Artemisina

Poecilosclerida

Ophlitaspongiinae

Poecilosclerida

Microcionidae

no

yes

Asbestopluma

Poecilosclerida

Cladorhizidae

Poecilosclerida

Cladorhizidae

no

yes

Asteropus

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

yes

Astrosclera

Agelasida

Astroscleridae

Agelasida

Astroscleridae

28S,COI,18S, ITS1-5.8S-ITS2-28S

no

Astrotylus

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Atergia

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

18S

no

Auletta

Halichondrida

Axinellidae

Bubarida

Bubaridae1

no

yes

Aulospongus

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Awhiowhio

‘Lithistida’

Corallistidae

Tetractinellida (Astrophorina)

Corallistidae

no

no

Axechina

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

18S, 28S, COI

no

Axinella

Halichondrida

Axinellidae

Axinellida

Axinellidae

18S, 28S

yes

Axinyssa

Halichondrida

Halichondriidae

Bubarida

Dictyonellidae

18S, 28S

yes

Axos

Hadromerida

Hemiasterellidae

Tethyida

Hemiasterellidae

28S, COI

yes

Baikalospongia

Haplosclerida

Lubomirskiidae

Spongillida

Lubomirskiidae

COI, cox2-ATP6, 18S, ITS1-5.8S-ITS2-28S

yes

Balliviaspongia

Haplosclerida

Spongillina incertae sedis

Spongillida

Spongillida incertae sedis

no

no

Batzella

Poecilosclerida

Chondropsidae

Poecilosclerida

Chondropsidae

no

no

Biemna

Poecilosclerida

Desmacelliae

Biemnida

Biemnidae

18S, 28S, COI

yes

Brachiaster

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

Pachastrellidae

no

no

Bubaris

Halichondrida

Bubaridae

Bubarida

Bubaridae

no

yes

Burtonitethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Callipelta

‘Lithistida’

Neopeltidae

Tetractinellida (Astrophorina)

Neopeltidae

no

yes

Calthropella

Astrophorida

Calthropellidae

Tetractinellida (Astrophorina)

Calthropellidae

no

yes

Caminella

Astrophorida

Geodiidae

Tetractinellida (Astrophorina)

Geodiidae

no

yes

Caminus

Astrophorida

Geodiidae

Tetractinellida (Astrophorina)

Geodiidae

COI

no

Cantabrina

Poecilosclerida

Echinodictyinae

Poecilosclerida

Echinodictyinae

no

no

Caulospongia

Hadromerida

Suberitidae

Suberitida

Suberitidae

no

yes

Celtodoryx

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

no (contaminant)

no

Ceratoporella

Agelasida

Astroscleridae

Agelasida

Astroscleridae

COI, ITS1-5.8S

no

Ceratopsion

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Cerbaris

Halichondrida

Bubaridae

Bubarida

Bubaridae

no

no

Cercicladia

Poecilosclerida

Cladorhizidae

Poecilosclerida

Cladorhizidae

no

no

Cervicornia

Hadromerida

Clionaidae

Clionaida

Clionaidae

18S, 28S

no

Chaetodoryx

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

no

no

Characella

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

(Astrophorina) incertae sedis

no

yes

Chelotropella

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

no

Chondrocladia

Poecilosclerida

Cladorhizidae

Poecilosclerida

Cladorhizidae

no

yes

Chondropsis

Poecilosclerida

Chondropsidae

Poecilosclerida

Chondropsidae

no

yes

Cinachyra

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

COI, 28S, 18S

yes

Cinachyrella

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

no

yes

Ciocalapata

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

no

no

Ciocalypta

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

18S, 28S, COI

yes

Cladorhiza

Poecilosclerida

Cladorhizidae

Poecilosclerida

Cladorhizidae

no

yes

Cladothenea

Astrophorida

Theneidae

Tetractinellida (Astrophorina)

Theneidae

no

no

Clathria

Poecilosclerida

Microcioninae

Poecilosclerida

Microcionidae

no

yes

Cliona

Hadromerida

Clionaidae

Clionaida

Clionaidae

18S, 28S, COI

yes

Clionaopsis

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

no

Cliothosa

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

no

Coelocarteria

Poecilosclerida

Isodictyidae

Poecilosclerida

Isodictyidae

28S

no

Coelodischela

Poecilosclerida

Guitarridae

Poecilosclerida

Guitarridae

no

no

Coelosphaera

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

no

no

Collectella

‘Lithistida’

Lithistida incertae sedis

Tetractinellida (Astrophorina)

Theonellidae

no

no

Columnitis

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Corallistes

‘Lithistida’

Corallistidae

Tetractinellida (Astrophorina)

Corallistidae

no

yes

Cornulella

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

yes

Cornulum

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Cortispongilla

Haplosclerida

Malawispongiidae

Spongillida

Malawispongiidae

COI, 18S, 5.8S-ITS2-28S

no

Corvoheteromeyenia

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Corvomeyenia

Haplosclerida

Metaniidae

Spongillida

Metaniidae

no

yes

Corvospongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Costifer

‘Lithistida’

Isoraphiniidae

Tetractinellida (Astrophorina)

Isoraphiniidae

no

no

Crambe

Poecilosclerida

Crambeidae

Poecilosclerida

Crambeidae

18S, 28S

yes

Craniella

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

no

yes

Crella

Poecilosclerida

Crellidae

Poecilosclerida

Crellidae

18S, 28S, tpm

yes

Crellastrina

Poecilosclerida

Crellidae

Poecilosclerida

Crellidae

no

no

Crellomima

Poecilosclerida

Crellidae

Poecilosclerida

Crellidae

no

no

Cryptax

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

no

no

Cryptosyringa

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

no

Cyamon

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

no

Cyclacanthia

Poecilosclerida

Latrunculiidae

Poecilosclerida

Latrunculiidae

no

no

Cymbastela

Halichondrida

Axinellidae

Bubarida

Dictyonellidae

28S

yes

Daedalopelta

‘Lithistida’

Neopeltidae

Tetractinellida (Astrophorina)

Neopeltidae

no

no

Damiria

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Damiriopsis

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

no

no

Delectona

Astrophorida

Thoosidae

Tetractinellida (Astrophorina)

Thoosidae

no

no

Dendoricella

Poecilosclerida

Dendoricellidae

Poecilosclerida

Dendoricellidae

no

no

Dercitus

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

COI, 18S

yes

Desmacella

Poecilosclerida

Desmacellidae

Desmacellida

Desmacellidae

18S, 28S

yes

Desmacidon

Poecilosclerida

Desmacididae

Poecilosclerida

Desmacididae

no

no

Desmanthus

‘Lithistida’

Desmanthidae

Bubarida

Bubaridae

18S, 28S, COI

no

Desmapsamma

Poecilosclerida

Desmacididae

Poecilosclerida

Desmacididae

16S, 18S, 28S, COI

no

Desmascula

‘Lithistida’

Azoricidae

Tetractinellida (Spirophorina)

Azoricidae

no

no

Desmoxya

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Diacarnus

Poecilosclerida

Podospongiidae

Poecilosclerida

Podospongiidae3

18S, 28S, COI

no

Dictyonella

Halichondrida

Dictyonellidae

Bubarida

Dictyonellidae

18S, 28S

yes

Didiscus

Halichondrida

Heteroxyidae

Axinellida

Raspailiidae

no

yes

Diplastrella

Hadromerida

Spirastrellidae

Clionaida

Spirastrellidae

no

yes

Diplopodospongia

Poecilosclerida

Podospongiidae

Poecilosclerida

Podospongiidae

no

no

Discodermia

‘Lithistida’

Theonellidae

Tetractinellida (Astrophorina)

Theonellidae

no

yes

Discorhabdella

Poecilosclerida

Crambeidae

Poecilosclerida

Crambeidae

no

no

Disyringa

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

18S, 28S

no

Dolichacantha

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Dosilia

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Dotona

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

no

Dragmacidon

Halichondrida

Axinellidae

Axinellida

Axinellidae

no

yes

Dragmatella

Poecilosclerida

Desmacellidae

Desmacellida

Desmacellidae

no

no

Dragmaxia

Halichondrida

Axinellidae

Axinellida

Axinellidae

no

yes

Drulia

Haplosclerida

Metaniidae

Spongillida

Metaniidae

5.8S-ITS2-28S

no

Duosclera

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Dyscliona

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

no

Echinochalina

Poecilosclerida

Microcioninae

Poecilosclerida

Microcionidae

no

yes

Echinoclathria

Poecilosclerida

Ophlitaspongiinae

Poecilosclerida

Microcionidae

no

yes

Echinodictyum

Poecilosclerida

Echinodictyinae

Poecilosclerida

Echinodictyinae

no

yes

Echinospongilla

Haplosclerida

Potamolepidae

Spongillida

Potamolepidae

COI, 18S, ITS1-5.8S-ITS2-28S

no

Echinostylinos

Poecilosclerida

Phellodermidae

Poecilosclerida

Phellodermidae

no

no

Ecionemia

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

18S

yes

Ectyonopsis

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

no

yes

Ectyoplasia

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

18S, 28S

yes

Endectyon

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Eospongilla

Haplosclerida

Palaeospongillidae

Spongillida

Palaeospongillidae

no

no

Ephydatia

Haplosclerida

Spongillidae

Spongillida

Spongillidae

COI, 18S, 5.8S-ITS2-28S, mt, ESTs

yes

Epipolasis

Halichondrida

Axinellidae

Axinellida

Axinellidae

no

yes

Erylus

Astrophorida

Geodiidae

Tetractinellida (Astrophorina)

Geodiidae

COI, 28S

yes

Esperiopsis

Poecilosclerida

Esperiopsidae

Poecilosclerida

Esperiopsidae

no

no

Euchelipluma

Poecilosclerida

Guitarridae

Poecilosclerida

Guitarridae

no

no

Eunapius

Haplosclerida

Spongillidae

Spongillida

Spongillidae

COI, ITS2, 18S

yes

Eurypon

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Exsuperantia

‘Lithistida’

Phymaraphiniidae

Tetractinellida (Astrophorina)

Phymaraphiniidae

no

yes

Fangophilina

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

no

yes

Fibulia

Poecilosclerida

Dendoricellidae

Poecilosclerida

Dendoricellidae

no

no

Forcepia

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

no

yes

Gastrophanella

‘Lithistida’

Siphonidiidae

Tetractinellida (Spirophorina)

Siphonidiidae

no

no

Geodia

Astrophorida

Geodiidae

Tetractinellida (Astrophorina)

Geodiidae

no

yes

Goreauiella

Agelasida

Astroscleridae

Agelasida

Astroscleridae

no

no

Guitarra

Poecilosclerida

Guitarridae

Poecilosclerida

Guitarridae

28S

no

Halichondria

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

18S, 28S, 5.8S, COI

yes

Halicnemia

Halichondrida

Heteroxyidae

Axinellida

Stelligeridae

no

yes

Halicometes

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Hamacantha

Poecilosclerida

Hamacanthidae

Merliida

Hamacanthidae

no

no

Hamigera

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

18S, 28S

yes

Hemiasterella

Hadromerida

Hemiasterellidae

Tethyida

Hemiasterellidae

no

yes

Hemimycale

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

18S, 28S

no

Hemitedania

Poecilosclerida

Tedaniidae

Poecilosclerida

Tedaniidae

no

no

Herengeria

‘Lithistida’

Corallistidae

Tetractinellida (Astrophorina)

Corallistidae

no

no

Heteromeyenia

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Heterorotula

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

yes

Heteroxya

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Higginsia

Halichondrida

Heteroxyidae

Axinellida

Stelligeridae

no

yes

Hispidopetra

Agelasida

Astroscleridae

Agelasida

Astroscleridae

no

no

Histodermella

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

no

no

Holopsamma

Poecilosclerida

Microcioninae

Poecilosclerida

Microcionidae

no

yes

Holoxea

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

yes

Homaxinella

Hadromerida

Suberitidae

Suberitida

Suberitida incertae sedis

no

yes

Homophymia

‘Lithistida’

Neopeltidae

Tetractinellida (Astrophorina)

Neopeltidae

no

yes

Houssayella

Haplosclerida

Metaniidae

Spongillida

Metaniidae

no

no

Hymedesmia

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

no

yes

Hymenancora

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

no

no

Hymeniacidon

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

18S, 28S, COI

yes

Hymeraphia

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

18S, 28S

yes

Hymerhabdia

Agelasida

Hymerhabdiidae

Agelasida

Hymerhabdiidae

18S, 28S, COI

no

Hymetrochota

Poecilosclerida

Iotrochotidae

Poecilosclerida

Iotrochotidae

no

no

Inflatella

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

18S

no

Iophon

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

yes

Iotroata

Poecilosclerida

Iotrochotidae

Poecilosclerida

Iotrochotidae

no

no

Iotrochopsamma

Poecilosclerida

Iotrochotidae

Poecilosclerida

Iotrochotidae

no

no

Iotrochota

Poecilosclerida

Iotrochotidae

Poecilosclerida

Iotrochotidae

18S, 28S, mt

no

Isabella

‘Lithistida’

Corallistidae

Tetractinellida (Astrophorina)

Corallistidae

no

no

Isodictya

Poecilosclerida

Isodictyidae

Poecilosclerida

Isodictyidae

no

yes

Janulum

Poecilosclerida

Raspailiidae

Haplosclerida

Phloeodictyidae

18S

no

Jaspis

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

yes

Jereicopsis

‘Lithistida’

Azoricidae

Tetractinellida (Spirophorina)

Azoricidae

no

no

Johannesia

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

no

no

Julavis

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Kaliapsis

‘Lithistida’

Phymaraphiniidae

Tetractinellida (Astrophorina)

Phymaraphiniidae

no

no

Kirkpatrickia

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

no

no

Lamellomorpha

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

(Astrophorina) incertae sedis

18S

no

Laminospongia

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

no

no

Latrunculia

Poecilosclerida

Latrunculiidae

Poecilosclerida

Latrunculiidae

no

yes

Laxotethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

28S

no

Leiodermatium

‘Lithistida’

Azoricidae

Tetractinellida (Spirophorina)

Azoricidae

no

no

Lepidosphaera

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

no

no

Lepidothenea

‘Lithistida’

Phymaraphiniidae

Tetractinellida (Astrophorina)

Phymaraphiniidae

no

no

Leptosastra

Hadromerida

Hemiasterellidae

Tethyida

Hemiasterellidae

no

no

Liosina

Halichondrida

Dictyonellidae

Tethyida

Hemiasterellidae

28S, COI

no

Lipastrotethya

Halichondrida

Dictyonellidae

Bubarida

Dictyonellidae

no

no

Lissodendoryx

Poecilosclerida

Coelosphaeridae

Poecilosclerida

Coelosphaeridae

no

yes

Lithobactrum

‘Lithistida’

Siphonidiidae

Tetractinellida (Spirophorina)

Siphonidiidae

no

no

Lithochela

Poecilosclerida

Crambeidae

Poecilosclerida

Crambeidae

no

no

Lithoplocamia

Poecilosclerida

Raspailiidae

Poecilosclerida

Raspailiidae

no

no

Lollipocladia

Poecilosclerida

Cladorhizidae

Poecilosclerida

Cladorhizidae

no

no

Lubomirskia

Haplosclerida

Lubomirskiidae

Spongillida

Lubomirskiidae

65 sequences, mt, ESTs

yes

Lutetiospongilla

Haplosclerida

Palaeospongillidae

Spongillida

Palaeospongillidae

no

no

Macandrewia

‘Lithistida’

Macandrewiidae

Tetractinellida (Astrophorina)

Macandrewiidae

no

no

Makedia

Haplosclerida

Spongillina incertae sedis

Spongillida

Spongillida incertae sedis

no

no

Malawispongia

Haplosclerida

Malawispongiidae

Spongillida

Malawispongiidae

no

no

Manihinea

‘Lithistida’

Theonellidae

Tetractinellida (Astrophorina)

Theonellidae

18S

no

Megaciella

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Melonanchora

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

no

no

Melophlus

Astrophorida

Geodiidae

Tetractinellida (Astrophorina)

Geodiidae

no

yes

Merlia

Poecilosclerida

Merliidae

Merliida

Merliidae

16S, 18S, 28S

no

Metania

Haplosclerida

Metaniidae

Spongillida

Metaniidae

no

no

Metschnikowia

Haplosclerida

Metschnikowiidae

Spongillida

Metschnikowiidae

no

no

Microscleroderma

‘Lithistida’

Scleritodermidae

Tetractinellida (Spirophorina)

Scleritodermidae

no

yes

Microtylostylifer

Poecilosclerida

Desmacellidae

Desmacellida

Desmacellidae

no

no

Microxistyla

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Monanchora

Poecilosclerida

Crambeidae

Poecilosclerida

Crambeidae

28S, COI

yes

Monocrepidium

Halichondrida

Bubaridae

Bubarida

Bubaridae

no

no

Mycale

Poecilosclerida

Mycalidae

Poecilosclerida

Mycalidae

no

yes

Myrmekioderma

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

18S, 28S, COI

yes

Myxilla

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

28S

yes

Myxodoryx

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

no

no

Neamphius

Astrophorida

Thoosidae

Tetractinellida (Astrophorina)

(Astrophorina) incertae sedis

COI, 28S

yes

Negombata

Poecilosclerida

Podospongiidae

Poecilosclerida

Podospongiidae

18S, COI

yes

Negombo

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Neoaulaxinia

‘Lithistida’

Phymatellidae

Tetractinellida (Astrophorina)

Phymatellidae

no

no

Neocladia

Poecilosclerida

Cladorhizidae

Poecilosclerida

Cladorhizidae

no

yes

Neofibularia

Poecilosclerida

Desmacellidae

Biemnida

Biemnidae

18S, 28S, COI

yes

Neopelta

‘Lithistida’

Neopeltidae

Tetractinellida (Astrophorina)

Neopeltidae

no

no

Neophrissospongia

‘Lithistida’

Corallistidae

Tetractinellida (Astrophorina)

Corallistidae

no

yes

Neopodospongia

Poecilosclerida

Podospongiidae

Poecilosclerida

Podospongiidae

no

yes

Neoschrammeniella

‘Lithistida’

Corallistidae

Tetractinellida (Astrophorina)

Corallistidae

no

no

Neosiphonia

‘Lithistida’

Phymatellidae

Tetractinellida (Astrophorina)

Phymatellidae

no

no

Nethea

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

Pachastrellidae

no

yes

Nucleotethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Nudospongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

yes

Ochridaspongia

Haplosclerida

Malawispongiidae

Spongillida

Malawispongiidae

no

no

Ohridospongilla

Haplosclerida

Spongillina incertae sedis

Spongillida

Spongillida incertae sedis

no

yes

Oncosclera

Haplosclerida

Potamolepidae

Spongillida

Potamolepidae

no

yes

Onotoa

Hadromerida

Placospongiidae

Clionaida

Placospongiidae

no

no

Ophiraphidites

Halichondrida

Axinellidae

Bubarida

Bubaridae

no

no

Ophlitaspongia

Poecilosclerida

Ophlitaspongiinae

Poecilosclerida

Microcionidae

16S, 18S

no

Oxytethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Pachastrella

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

Pachastrellidae

no

yes

Pachydictyum

Haplosclerida

Malawispongiidae

Spongillida

Malawispongiidae

COI, 18S, ESTs

yes

Pachymatisma

Astrophorida

Geodiidae

Tetractinellida (Astrophorina)

Geodiidae

COI, 28S, 18S, ITS1-5.8S-ITS2

yes

Pachyrotula

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Palaeospongilla

Haplosclerida

Palaeospongillidae

Spongillida

Palaeospongillidae

no

no

Pandaros

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

COI

no

Paracornulum

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

COI

no

Paradesmanthus

‘Lithistida’

Desmanthidae

Bubarida

Bubaridae

no

no

Parahigginsia

Halichondrida

Heteroxyidae

Axinellida

Heteroxyidae

no

no

Pararhaphoxya

Halichondrida

Axinellidae

Bubarida

Bubaridae

no

yes

Paratetilla

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

no

yes

Paratimea

Hadromerida

Stelligeridae

Axinellida

Stelligeridae

no

yes

Pectispongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Penares

Astrophorida

Geodiidae

Tetractinellida (Astrophorina)

Geodiidae

5.8S-ITS2-28S

yes

Penicillus

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Petromica

‘Lithistida’

Desmanthidae

incertae sedis

 

no

yes

Phakellia

Halichondrida

Axinellidae

Bubarida

Bubaridae

18S, 28S, COI

yes

Phakettia

Halichondrida

Dictyonellidae

Bubarida

Dictyonellidae

no

yes

Phelloderma

Poecilosclerida

Phellodermidae

Poecilosclerida

Phellodermidae

no

yes

Phlyctaenopora

Poecilosclerida

Mycalidae

Poecilosclerida

Mycalidae

no

no

Phorbas

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

18S

yes

Phoriospongia

Poecilosclerida

Chondropsidae

Poecilosclerida

Chondropsidae

no

no

Phycopsis

Halichondrida

Axinellidae

Bubarida

Bubaridae

no

yes

Pione

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

yes

Pipestela

Halichondrida

Axinellidae

Bubarida

Bubaridae

no

no

Placospherastra

Hadromerida

Placospongiidae

Clionaida

Placospongiidae

no

no

Placospongia

Hadromerida

Placospongiidae

Clionaida

Placospongiidae

18S, COI

yes

Plakidium

‘Lithistida’

Lithistida incertae sedis

incertae sedis

 

no

no

Pleroma

‘Lithistida’

Pleromidae

Tetractinellida (Astrophorina)

Pleromidae

no

no

Plicatellopsis

Hadromerida

Suberitidae

Suberitida

Suberitidae

no

no

Plocamiancora

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

no

yes

Plocamione

Poecilosclerida

Raspailiidae

Poecilosclerida

Raspailiidae

no

no

Plocamionida

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

18S, 28S, COI

yes

Podospongia

Poecilosclerida

Podospongiidae

Poecilosclerida

Podospongiidae

no

no

Poecillastra

Astrophorida

Vulcanellidae

Tetractinellida (Astrophorina)

Vulcanellidae

COI, 28S

yes

Polymastia

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

yes

Pomelia

‘Lithistida’

Scleritodermidae

Tetractinellida (Spirophorina)

Scleritodermidae

no

no

Poritella

‘Lithistida’

Lithistida incertae sedis

incertae sedis

 

no

no

Potamolepis

Haplosclerida

Potamolepidae

Spongillida

Potamolepidae

no

yes

Potamophloios

Haplosclerida

Potamolepidae

Spongillida

Potamolepidae

no

no

Pottsiela

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Pozziella

Poecilosclerida

Hamacanthidae

Merliida

Hamacanthidae

no

no

Prosuberites

Hadromerida

Suberitidae

Agelasida

Hymerhabdiidae

28S

yes

Proteleia

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

18S

no

Protosuberites

Hadromerida

Suberitidae

Suberitida

Suberitidae

no

yes

Psammastra

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

no

Psammochela

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

no

no

Psammoclema

Poecilosclerida

Chondropsidae

Poecilosclerida

Chondropsidae

no

yes

Pseudohalichondria

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

no

no

Pseudospongosorites

Hadromerida

Suberitidae

Suberitida

Suberitidae

tpm

no

Pseudosuberites

Hadromerida

Suberitidae

Suberitida

Suberitidae

no

yes

Pseudotrachya

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Ptilocaulis

Halichondrida

Axinellidae

Axinellida

Raspailiidae

18S, 28S, mt

yes

Pyloderma

Poecilosclerida

Dendoricellidae

Poecilosclerida

Dendoricellidae

no

no

Quasillina

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

18S

no

Racekiela

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Racodiscula

‘Lithistida’

Theonellidae

Tetractinellida (Astrophorina)

Theonellidae

no

no

Radiella

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Radiospongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

yes

Raspaciona

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

18S, 28S, COI

no

Raspailia

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Reidispongia

‘Lithistida’

Phymatellidae

Tetractinellida (Astrophorina)

Phymatellidae

no

no

Reniochalina

Halichondrida

Axinellidae

Axinellida

Raspailiidae

18S, 28S

yes

Rezinkovia

Haplosclerida

Lubomirskiidae

Spongillida

Lubomirskiidae

18S-5.8S-ITS2-28S, mt

yes

Rhabdastrella

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Geodiidae

no

yes

Rhabderemia

Poecilosclerida

Rhabderemiidae

Biemnida

Rhabderemiidae

no

yes

Rhabdeurypon

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

no

Rhabdobaris

Halichondrida

Bubaridae

Bubarida

Bubaridae

no

no

Rhaphidhistia

Hadromerida

Trachycladidae

Trachycladida

Trachycladidae

no

no

Rhaphoxya

Halichondrida

Dictyonellidae

Bubarida

Dictyonellidae

no

no

Rhizaxinella

Hadromerida

Suberitidae

Suberitida

Suberitidae

no

yes

Ridleia

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Rotuloplocamia

Poecilosclerida

Iotrochotidae

Poecilosclerida

Iotrochotidae

no

no

Samus

Spirophorida

Samidae

Tetractinellida (Spirophorina)

Samidae

no

no

Sanidastra

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Sarcomella

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

no

no

Saturnospongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Sceptrella

Poecilosclerida

Latrunculiidae

Poecilosclerida

Latrunculiidae

no

yes

Sceptrintus

Poecilosclerida

Podospongiidae

Poecilosclerida

Podospongiidae

no

no

Scleritoderma

‘Lithistida’

Scleritodermidae

Tetractinellida (Spirophorina)

Scleritodermidae

no

no

Scolopes

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

no

Scopalina

Halichondrida

Scopalinidae

Scopalinida

Scopalinidae

18S, 28S, COI

yes

Semisuberites

Poecilosclerida

Esperiopsidae

Poecilosclerida

Esperiopsidae

no

no

Setidium

‘Lithistida’

Scleritodermidae

Tetractinellida (Spirophorina)

Scleritodermidae

no

no

Sigmaxinella

Poecilosclerida

Desmacellidae

Biemnida

Biemnidae

no

yes

Sigmeurypon

Poecilosclerida

Ophlitaspongiinae

Poecilosclerida

Microcionidae

no

no

Sigmosceptrella

Poecilosclerida

Podospongiidae

Poecilosclerida

Podospongiidae

no

no

Siliquariaspongia

‘Lithistida’

Theonellidae

Tetractinellida (Astrophorina)

Theonellidae

no

no

Siphonidium

‘Lithistida’

Siphonidiidae

Tetractinellida (Spirophorina)

Siphonidiidae

no

no

Sollasella

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Spanioplon

Poecilosclerida

Hymedesmiidae

Poecilosclerida

Hymedesmiidae

18S, 28S

no

Sphaerotylus

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

yes

Spheciospongia

Hadromerida

Clionaidae

Clionaida

Clionaidae

18S

yes

Spinospongilla

Haplosclerida

Malawispongiidae

Spongillida

Malawispongiidae

no

no

Spinularia

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Spirasigma

Spirophorida

Spirasigmidae

Tetractinellida (Spirophorina)

Spirasigmidae

no

no

Spirastrella

Hadromerida

Spirastrellidae

Clionaida

Spirastrellidae

no

yes

Spirorhabdia

Poecilosclerida

Crellidae

Poecilosclerida

Crellidae

no

no

Spiroxya

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

no

Spongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

COI,18S, 28S, ITS1-5.8S-ITS2-28S, tpm

yes

Spongosorites

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

no

yes

Stelletta

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

COI, 28S, 18S

yes

Stellettinopsis

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

yes

Stelligera

Hadromerida

Stelligeridae

Axinellida

Stelligeridae

18S, 28S, COI

yes

Stellitethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

yes

Stelodoryx

Poecilosclerida

Myxillidae

Poecilosclerida

Myxillidae

no

no

Sterrastrolepis

Haplosclerida

Potamolepidae

Spongillida

Potamolepidae

no

no

Stratospongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Stromatospongia

Agelasida

Astroscleridae

Agelasida

Astroscleridae

no

no

Strongylacidon

Poecilosclerida

Chondropsidae

Poecilosclerida

Chondropsidae

no

yes

Strongylamma

Poecilosclerida

Tedaniidae

Poecilosclerida

Tedaniidae

no

no

Strongylodesma

Poecilosclerida

Latrunculiidae

Poecilosclerida

Latrunculiidae

no

no

Stryphnus

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

yes

Stylissa

Halichondrida

Scopalinidae

Scopalinida

Scopalinidae

18S, 28S, COI

yes

Stylocordyla

Hadromerida

Stylocordylidae

Suberitida

Stylocordylidae

COI (submitted)

no

Suberites

Hadromerida

Suberitidae

Suberitida

Suberitidae

18S, 28S, mt, ESTs

yes

Sulcastrella

‘Lithistida’

Desmanthidae

incertae sedis

 

no

no

Svenzea

Halichondrida

Scopalinidae

Scopalinida

Scopalinidae

18S, 28S, COI

yes

Swartschewskia

Haplosclerida

Lubomirskiidae

Spongillida

Lubomirskiidae

COI, 18S, 5.8S-ITS2-28S, mt

no

Tectitethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

yes

Tedania

Poecilosclerida

Tedaniidae

Poecilosclerida

Tedaniidae

no

yes

Tedaniphorbas

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Tentorina

Spirophorida

Spirasigmidae

Tetractinellida (Spirophorina)

Spirasigmidae

no

no

Tentorium

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

18S

no

Terpios

Hadromerida

Suberitidae

Suberitida

Suberitida incertae sedis

no

yes

Tethya

Hadromerida

Tethyidae

Tethyida

Tethyidae

18S, 28S, COI

yes

Tethyastra

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Tethycometes

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

no

Tethyopsis

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

yes

Tethyspira

Halichondrida

Dictyonellidae

Axinellida

Raspailiidae

18S, 28S, COI

no

Tethytimea

Hadromerida

Tethyidae

Tethyida

Tethyidae

no

yes

Tetilla

Spirophorida

Tetillidae

Tetractinellida (Spirophorina)

Tetillidae

no

yes

Tetrapocillon

Poecilosclerida

Guitarridae

Poecilosclerida

Guitarridae

no

yes

Thenea

Astrophorida

Theneidae

Tetractinellida (Astrophorina)

Theneidae

COI, 28S

yes

Theonella

‘Lithistida’

Theonellidae

Tetractinellida (Astrophorina)

Theonellidae

COI, 28S, 18S

yes

Thoosa

Astrophorida

Thoosidae

Tetractinellida (Astrophorina)

Thoosidae

no

no

Thrinacophora

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Thrombus

Astrophorida

Thrombidae

Tetractinellida (Astrophorina)

Thrombidae

28S

no

Timea

Hadromerida

Timeidae

Tethyida

Tethyidae

no

yes

Topsentia

Halichondrida

Halichondriidae

Suberitida

Suberitida incertae sedis

no

yes

Trachostylea

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

no

Trachycladus

Hadromerida

Trachycladidae

Trachycladida

Trachycladidae

18S, 28S

yes

Trachyteleia

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Tribrachium

Astrophorida

Ancorinidae

Tetractinellida (Astrophorina)

Ancorinidae

no

no

Trikentrion

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

yes

Triptolemma

Astrophorida

Pachastrellidae

Tetractinellida (Astrophorina)

Pachastrellidae

no

yes

Trochospongilla

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

yes

Tsitsikamma

Poecilosclerida

Latrunculiidae

Poecilosclerida

Latrunculiidae

18S, COI

no

Tylexocladus

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Ulosa

Poecilosclerida

Esperiopsidae

Poecilosclerida2

Esperiopsidae2

no

yes

Umborotula

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Uritaia

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

no

no

Uruguaya

Haplosclerida

Potamolepidae

Spongillida

Potamolepidae

no

no

Uruguayella

Haplosclerida

Spongillidae

Spongillida

Spongillidae

no

no

Vetulina

‘Lithistida’

Vetulinidae

Sphaerocladina

Vetulinidae

18S

yes

Volzia

Hadromerida

Clionaidae

Clionaida

Clionaidae

no

no

Vosmaeria

Halichondrida

Halichondriidae

Suberitida

Halichondriidae

28S

no

Vulcanella

Astrophorida

Vulcanellidae

Tetractinellida (Astrophorina)

Vulcanellidae

no

yes

Waltherarndtia

Poecilosclerida

Raspailiidae

Axinellida

Raspailiidae

no

no

Weberella

Hadromerida

Polymastiidae

Polymastiida

Polymastiidae

no

no

Wigginsia

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

no

Willardia

Hadromerida

Acanthochaetetidae

Clionaida

Acanthochaetetidae

no

no

Xenospongia

Hadromerida

Tethyidae

Tethyida

Tethyidae

18S, 28S

no

Yucatania

Astrophorida

Thrombidae

Tetractinellida (Astrophorina)

Thrombidae

no

no

Zyzzya

Poecilosclerida

Acarnidae

Poecilosclerida

Acarnidae

no

yes

  1. 1 One sequence of Auletta in Erpenbeck et al. [39] groups with Acanthella and Phakellia ventilabrum.
  2. 2 28S sequences of Ulosa digitata suggest that this genus could belong to the Suberitida, as it groups with Halichondriidae [5].
  3. 3 Redmond et al. [10] show D. spinipoculum (type species) clustering with Raspailiidae whilst D. bismarckensis groups with Podospongiidae. It is possible that the 18S sequence of D. spinipoculum in Redmond et al. [10] is a result of contamination and therefore we retain Diacarnus in Podospongiidae.