Overview of left and right inner ears including otoliths in (A-B) S. tinanti , (C-D) H. guttatus , and (E-F) E. maculatus . Inner ears are shown in posterior (A, C, E) and dorsal views (B, D, F). While inner ears are rather close to each other in H. guttatus and E. maculatus, the distance between left and right ears is distinctly larger in S. tinanti. In E. maculatus the lagenae come very close to each other and the spatial orientation of the semicircular canals—especially of the horizontal semicircular canal—differ from the other two species. Note that semicircular canals of the left inner ear of E. maculatus are incomplete. The 3D reconstructions are based on microCT scans (isotropic voxel size: 10 μm). a, anterior; asc, anterior semicircular canal; cc, common canal; d, dorsal; hsc, horizontal semicircular canal; l, lateral; lag, lagena; lot, lagenar otolith; psc, posterior semicircular canal; sac, saccule; sot, saccular otolith; uot, utricular otolith; utr, utricle. Scale bar, 1 mm.